The first notosuchian crocodyliform from the Araçatuba Formation (Bauru Group, Paraná Basin), and diversification of sphagesaurians
André Eduardo Piacentini Pinheiro, Rafael Gomes de Souza, Kamila L. N. Bandeira, Arthur Souza Brum, Paulo Victor Luiz Gomes da Costa Pereira, Luís Otávio Rezende Castro, Renato Rodríguez Cabral Ramos, Felipe Medeiros Simbras
Abstract
The mesoeucrocodylian record from the Bauru Group (Paraná Basin, Brazil) comes from three formations: Adamantina (Late Turonian-Early Maastrichtian), Presidente Prudente (Early Campanian-Early Maastrichtian) and Marília (Maastrichtian). These records are restricted to the Early Coniacian-Early Maastrichtian. Here, we report a new crocodyliform record from the Bauru Group, which was the first of the Araçatuba Formation, from a new locality of Coronel Goulart district, in Álvares Machado municipality. Coronelsuchus civali gen. nov. et sp. nov. comprises two related specimens - FFP PG 13 and FFP PG 14 recovered close to each other and on the same muddy sandstone level. We perform a phylogenetic analysis with 392 characters (five new added) and 88 taxa. Our analyses supported the new clades: Sphagesauria, Sphagesaurinae and Caipirasuchinae. Our phylogenetic results suggest C. civali as a basal Sphagesauria. Regarding the Crocodyliform Assemblage Zones (CAZ), the new species is correlated to the CAZ 1, together with small sphagesaurians, Caryonosuchus and itasuchids. The C. civali reveals an older appearing (pre-Turonian) and diversification of Sphagesauria. The specimens also expand the stratigraphic range for Notosuchia in the Bauru Group.