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Plasma membrane H<sup>+</sup>‐ATPases‐mediated cytosolic proton gradient regulates pollen tube growth

Wei Chen, Pengfei Jia, Wei‐Cai Yang, Hong‐Ju Li

2020Journal of Integrative Plant Biology25 citationsDOI

Abstract

Abstract The polar growth of pollen tubes is essential for the delivery of sperm cells during fertilization in angiosperms. How this polar growth is regulated has been a long‐standing question. An in vitro pharmacological assay previously implicated proton flux in pollen tube growth, although genetic and cellular supporting evidence was lacking. Here, we report that protons form a gradient from the pollen tube tip to the shank region and this gradient is generated by three members of A rabidopsis H + ‐ A TPases (AHAs). Genetic analysis suggested that these AHAs are essential for pollen tube growth, thus providing new insight into the regulation of polar growth.

Topics & Concepts

Pollen tubePollenProtonGametophytePolarSpermBiologyATPaseElectrochemical gradientBiophysicsBotanyHuman fertilizationCell biologyProton pumpTube (container)MembraneBiochemistryPollinationGeneticsPhysicsEnzymeMaterials scienceAstronomyQuantum mechanicsComposite materialPhotosynthetic Processes and MechanismsPlant Reproductive BiologyPlant Molecular Biology Research
Plasma membrane H<sup>+</sup>‐ATPases‐mediated cytosolic proton gradient regulates pollen tube growth | Litcius